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Forms of
Algae
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Unicells:
single cells, motile or nonmotile
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Colonies:
Assemblage of individual cells with variable or constant number of cells
that remain constant throughout the colony life
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Coenobium:
Colony with constant number of cells, which cannot survive alone; specific
„tasks“ among groups of cells is common
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Filaments:
daughter cells remain attached after cell division and form a cell chain;
adjacent cells share cell wall (distinguish them from linear colonies!);
maybe unbranched (uniseriate) or branched (mutiseriate)
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Coenocytic
or siphonaceaous forms: one large, multinucleate
cell without cross walls
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Parenchymatous
and pseudoparenchymatous algae: mostly macro-scopic
algae with tissue of undifferentiated cells and growth originating from
a meristem with cell division in three dimensions; pseudoparenchymatous
superficially ressemble parenchyma but are composed of appressed filaments
Phylogeny of Algae
-
Algae
did not evolve from one single, common ancestor but evolved at different
times from different ancestors.
-
Algae
are, therefore, not monophyletic
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Some
algae are closer related to heterotrophic flagellates or ciliates than
to other algae
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Prokaryotic
phototrophs: Cyanobacteria („blue-green algae“)
-
Eukaryotic
algae: possess a doublemembrane enclosed nucleus and ususal other membrane-enclosed
organelles (mitochondria, chloroplasts)
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Origin
of organelles: symbiotic enclosure of free living bacteria or cyanobacteria,
horizontal transfer of genes from endosymbionts to host nucleus
Nutrition of Algae
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Photosynthesis
is the primary mode of nutrition (C uptake as CO2)
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Photosynthetic
algae were mostly considered photoautotrophic
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Numerous
phototrophic algae can, however, take up dissolved organic matter or engulf
bacteria and other algal cells as particulate prey; they are referred to
as mixotrophic
-
Uptake
of particulate prey = phagotrophy;Uptake
of dissolved substances = osmotrophy
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The degree
of mixotrophie in chlorophyll-containing algae varies with environmental
conditions (light, prey abundance)
-
Some
chlorophyll-containing species can even lose their pigments when living
heterotrophic
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Most
algal phyla/divisions contain mixo- or heterotrophic species
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Some
algae cannot synthesize essential vitamins (biotin, thiamine, cobalamin
= B12) and have to import them: auxotrophic
species
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